Microsaccades uncover the orientation of covert attention
Our eyes perform small movements even while we look at stationary
visual scene. These fixational eye movements are subdivided
into tremor, drift, and microsaccades. Microsaccades are the fastest
components of fixational eye movements. Here, we investigated the
statistics of microsaccades in a classical spatial cuing paradigm
(Posner, 1980). First, we reproduced microsaccade suppression
with a minimum rate about 150 ms after cue onset - a well-known effect
reported first by Findlay (1976), Winterson & Collewijn (1976), and
Bridgeman & Palca (1980). Second, as a new finding we observe microsaccadic
enhancement with a maximum rate about 350 ms after presentation of
the cue. Third, we find a modulation of the orientation towards
the cue direction. Therefore, attentional influences can bias the
unconscious motor behavior during visual fixation. These results
suggest that microsaccades can be exploited to map the orientation of
visual attention in psychophysical experiments.
Animation: Fixational eye movements and microsaccades. In a simple
fixation task rather erratic miniature eye movements are observed. The
example
represents a fixation with a duration of 2348 ms (or 588 data samples),
recorded binocularly with an SMI EyeLink System (250 Hz). Microsaccades
are small but rapid events which can be identified as approximately
linear epoches of the eyes' trajectory (left eye: green, right eye:
red). The original size of the fixation cross shown in the animation is
0.73°. Data are the same as in Figure 1 of the above paper.
(Created by Jochen
Laubrock.)
Methods: Detection of microsaccades
Microsaccades are detected in 2D velocity space. The detection
thresholds are computed via a median-based estimation of the standard
deviations of horizontal and vertical components of the eyes's
velocity. We use a temporal overlap criterion to test for binocular
microsaccades. This procedure further reduces noise in the
detection algorithm. A MATLAB
implementation of the algorithm with a short sequence of experimental
data can be downloaded using the following files:
- xdemo.m
(executable main program)
- microsacc.m
(detection of microsaccades based on monocular data)
- binsacc.m
(testing for binocular microsaccades)
- vecvel.m
(compute 2D velocity vectors from position data)
- demo.dat
(raw data for running the demo)